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    <Title>EMODnet Biology GeoServer - WMS</Title>
    <Abstract>The EMODnet Biology products include a set of gridded map layers showing the average abundance of marine species for different time windows (seasonal, annual) using geospatial modelling. The spatial modelling tool used to calculate the gridded abundance maps is based on DIVA. DIVA (Data-Interpolating Variational Analysis) is a tool to create gridded data sets from discrete point measurements of the ocean. For the representation of time dynamics, it was decided to produce gridded maps for sliding time windows.</Abstract>
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        <ContactOrganization>Flanders Marine Institute (VLIZ)</ContactOrganization>
      </ContactPersonPrimary>
      <ContactPosition/>
      <ContactAddress>
        <AddressType/>
        <Address/>
        <City>Oostende</City>
        <StateOrProvince/>
        <PostCode>8400</PostCode>
        <Country>Belgium</Country>
      </ContactAddress>
      <ContactVoiceTelephone/>
      <ContactFacsimileTelephone/>
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      <Title>EMODnet Biology GeoServer - WMS</Title>
      <Abstract>The EMODnet Biology products include a set of gridded map layers showing the average abundance of marine species for different time windows (seasonal, annual) using geospatial modelling. The spatial modelling tool used to calculate the gridded abundance maps is based on DIVA. DIVA (Data-Interpolating Variational Analysis) is a tool to create gridded data sets from discrete point measurements of the ocean. For the representation of time dynamics, it was decided to produce gridded maps for sliding time windows.</Abstract>
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        <Name>abundance_benthos_North_Baltic_Seas</Name>
        <Title>Data product numerical abundance of benthic macroinvertebrates in North Sea and Baltic Sea</Title>
        <Abstract>The data product on benthic living modes (Beauchard, 2018), was based on an extensive compilation of data on benthic abundance and biomass. However, this dataset was only present as a data file, without the underlying scripts to reproduce the result. With the present data product, we correct this procedural gap. This dataset differs in details from the file underlying the data product on living modes. Datasets were selected that were sufficiently similar in methods for sampling (either boxcore or grab), sampled surface (in the order of 0.1 square meter, although the exact value is variable - it can be found back in the data files) and sieves (1 mm and 0.5 mm sieves were included). For all datasets, abundance was either used directly from the given abundance in the dataset, or calculated from the given counts and area sampled.</Abstract>
        <KeywordList>
          <Keyword>abundance</Keyword>
          <Keyword>WCS</Keyword>
          <Keyword>NetCDF</Keyword>
        </KeywordList>
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        <Abstract>Mixoplankton (sensu Flynn et al., 2019) is a newly introduced term indicating plankton that is capabable of both photosynthesis and phagotrophy. More details are found in Flynn et al., (2019). The potential trophic state can be seen as an inherent characteristic of plankton species. A literature and expert-knowledge study has provided the classification in either phototrophy or mixotrophy which is submitted as traits data to WoRMS. This analysis makes use of this classification to estimate the spatial and temporal distribution of the fraction of mixoplanktonic species in the Greater North Sea including the Celtic Seas. This project has received funding from the European Union’s Horizon 2020 research and innovation programme under the Marie Skłodowska-Curie grant agreement No 766327.</Abstract>
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        <Title>Habitat suitability of reef-froming species in the North Sea</Title>
        <Abstract>Due to fishing and other human activities, reef forming species have almost completely disappeared over roughly the past century. They are important structures that accomodate juvenile fish and other small organisms. For protection of areas where such reefs could possibly be reintroduced, it is important to define areas that are suitable habitats. This product aims to classify areas in the North Sea based on current occurence in combination with environmental variables that are particularly suitable for these organisms.</Abstract>
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        <Title>Habitat suitability maps for harbour porpoises in the North Sea-Baltic Sea transition zone from 1997 to 2022</Title>
        <Abstract>The dataset includes 6  layers showing modelled habitat suitability for harbour porpoises (Phocoena phocoena) across the North Sea–Baltic Sea transition zone over a 25-year period (1997–2004; 2005-2012; 2013-2022) during winter (Nov–Apr) and summer (May–Oct). Habitat suitability was estimated using the maximum entropy (MaxEnt) algorithm, based on satellite tag data of 111 individuals and a range of dynamic and static environmental conditions as predictor variables. Habitat suitability models and maps were developed by researchers of the Department of Ecoscience, Aarhus University to quantify potential spatiotemporal changes in the overlap between suitable habitats and an existing network of 62 MPAs dedicated to harbour porpoise protection. The habitat suitability rasters can inform effective design of forthcoming MPAs under the EU Biodiversity Strategy for 2030.&#13;
Harbour porpoise habitat suitability layers cover a seascape area of 70,145 km2. Habitat suitability layers are available for the periods: 1997–2004; 2005-2012; 2013-2022 and for both winter (Nov–Apr) and summer (May–Oct). All layers are projected using the EEA reference grid (ETRS89-LAEA Europe, EPSG:3035), have a 1x1 km spatial resolution (cell size) and a spatial extent of 4275000, 4545000, 3415000, 4105000  (xmin, xmax, ymin, ymax). Habitat suitability values (probabilities) range between 1 and 0 with values closer to 1 indicating habitat being highly suitable for harbour porpoises while values closer to 0 indicate habitat that is poorly suitable for harbour porpoises. Location data underlying the habitat suitability models were collected from 111 harbour porpoises tracked over the period 1997–2022 as part of a long-term satellite telemetry monitoring program in Denmark. Location and meta data are freely available at Zenodo (https://doi.org/10.5281/zenodo.13968991). &#13;
Harbour porpoise habitat suitability layers were produced as part of the projects: GES4SEAS (grant agreement no. 101059877) and OBAMA-NEXT (101091642))</Abstract>
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        <Title>Global predicted thermal distributions for intertidal macroalgae</Title>
        <Abstract>Ocean warming is expected to drive quick shifts in marine species distributions modifying coastal communities. Previous research has shown how the southern Bay of Biscay is particularly affected and the canopy-forming subtidal and intertidal macroalgal assemblages are expected to be replaced by turf-forming Mediterranean-like communities with warmer affinities by the end of the century. The effect of temperature on macroalgal communities could be assessed using thermal metrics that can target temperature effects alone. An example is the Community Temperature Index (CTI) which is an abundance-weighted mean of the species’ optimal temperature. It has been previously used to understand the thermal preferences of different organisms such us birds, fish or invertebrates.</Abstract>
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        <Title>Modelled projections of habitat for commercial fish around North-western Europe under climate change, 2020 to 2060</Title>
        <Abstract>Environmental Niche Model (ENM) outputs for 49 commercial fish species under climate change until the decade of 2060 around northwestern Europe. A model ensemble of 5 ENMs was used (MaxEnt, Generalised Linear Models, Support Vector Machine, Random Forest and BIOCLIM ), and projections were made under three different emission scenarios: A1B, RCP4.5 and RCP 8.5. The data shows model agreement (normalised to 1) for presence/absence decadal projections from 2020 to 2060. Additionally we provide data on model performance, with the Area Under the Curve (AUC) scores of the Receiver Operator Characteristic (ROC) curve for each of the 5 ENMs trained for each combination of fish species and emission scenario. Only ENMs with an AUC score of at least 0.7 were considered.&#13;
The data is based on the output of an ensemble of 5 Environmental Niche Modeling techniques: Maximum Entropy (MaxEnt), Generalised Linear Model (GLM), Support Vector Machine (SVM), Random Forest and BIOCLIM. Each model was trained to capture the environmental requirements of individual fish species, using the 20 year-average in the period 1997-2016 of the environmental variables: near-bed sea temperature, sea surface temperature, near-bed salinity, sea surface salinity, the difference in near-bed and surface salinity and temperature (an approximation of stratification), and depth.&#13;
Once the models were trained, they were used to study future habitat suitability for the 49 commercially-important fish species in 20-year averages centered in 2020, 2030, 2040, 2050 and 2060. To produce maps of the suitable habitat, the model projections were converted to binary presence/absence data using the thresholds that optimized the True Skill Statistics (TSS) of each model. The data shows the model agreement, normalised to 1 ("1" denotes all ENMs agree an area is suitable, "0" denotes all ENMs agree an area is not suitable). The number of ENMs considered for each combination of fish species and emission scenario depends on model performance. A ENM model was considered if its Area Under the Curve (AUC) score of the Receiver Operator Characteristic (ROC) curve was at least 0.7. Model performance is listed in the file "Model_performance_AUC.xlsx".&#13;
Model input data: Climate change projection data from Met Office and Plymouth Marine Laboratory (PML) and species presence and abundance records from online databases (Ocean Biodiversity Information System (OBIS), Global Biodiversity Information System (GBIF) and Marine Scotland (Moriarty, M., Greenstreet, S.P.R. and Rasmussen, J. (2017) Derivation of Groundfish Survey Monitoring and Assessment Data Product for the Northeast Atlantic Area. Scottish Marine and Freshwater Science, 8, 240pp. DOI: 10.7489/1984-1). Models were trained using the R and Maxent softwares. &#13;
Study region: northeast Atlantic shelf. Latitudes between -17° and 9.25°; longitudes between 44° and 65°.&#13;
These datasets were produced for the study in the article "Climate change projections of commercial fish distribution and suitable habitat around northwestern Europe", by Bryony L. Townhill, Elena Couce, Jonathan Tinker, Susan Kay, John K. Pinnegar, in Fish and Fisheries (2023; http://doi.org/10.1111/faf.12773).</Abstract>
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        <Title>Modelled projections of habitat for fish species feeding guilds around North-western Europe under climate change, 2010 to 2095</Title>
        <Abstract>These data are Bayesian Additive Regression Tree model annual predictions for habitat suitability of marine fish species across a range of body sizes and belonging to different feeding guilds from 2010 to 2095 in 5 year intervals in the northeast Atlantic shelf seas. Feeding guilds were allocated based on classifications following Thompson et al. (2020).&#13;
The purpose of this study was to predict how climate change could affect the species richness of consumers lower in the food web (planktivores), compared to those intermediate (benthivores) and higher up (piscivores).  In brief, a collation of trophic interactions spanning the northeast Atlantic shelf seas has been applied to define feeding guilds by grouping fish species size classes (we use the taxonomic level of Gobiidae and Ammodytes for taxa that are not consistently identified to species) that have prey taxa in common, and whose prey differentiate them from other predator guilds based on cluster analysis. Taxon-specific size categories were defined as: &lt;3 cm as larvae; small juvenile fish between 3 cm and half of length at maturity; juvenile-medium fish from half of length at maturity to length at maturity; medium fish from length at maturity to half-maximum length; and all remaining larger fish as large. Taxon-specific length at maturity and maximum length (i.e., asymptotic length at infinity) were estimated using the R package Fishlife (Thorson et al., 2017).&#13;
Environmental projections were derived from the coupled marine ecosystem models POLCOMS/NEMO-ERSEM. We focus on two emission scenarios (representative concentration pathways, RCPs) developed for the Intergovernmental Panel on Climate Change (IPCC)’s fifth phase of the Coupled Model Intercomparison Project (CMIP5): RCP4.5, the “stabilization scenario”, characterised by medium emissions and high mitigations, and the “no mitigation policy” scenario RCP8.5, derived from high fossil fuel emission and low mitigations. We use projections of temperature, salinity, pH, nitrate, phosphate, dissolved oxygen, current velocity, chlorophyll, gross primary production, non-living organic carbon, zooplankton carbon concentration and secondary carbon production by zooplankton in our predictive habitat models. For temperature, salinity, current velocity, dissolved oxygen, and pH both surface and bottom mean annual averages were considered, and in the case of temperature, also the difference between bottom and surface values, to account for stratification. For chlorophyll, gross primary production, non-living organic carbon, phosphate, nitrate, zooplankton carbon concentration and secondary carbon production by zooplankton the total across the water column was used, rather than surface or bottom values. We also include depth (from the General Bathymetric Chart of the Oceans GEBCO; www.gebco.net, at 15 second resolution), distance to coast and substrate composition (median grain size and percentages of mud, sand and gravel from Wilson et al., 2018) to capture key spatial gradients that affect habitat suitability for fish. &#13;
All environmental data was processed onto a 10 km by 10 km grid, and because an annual mean of, e.g., temperature, does not capture the environmental variability that ultimately determines the thresholds within which biota must survive; for temperature, salinity, pH, oxygen and current speed we also include the standard deviation of the 12 monthly means in each year, for all locations within a radius of 75 km of each grid cell, in order to provide a measure of spatio-temporal heterogeneity. For the environmental variables where surface and bottom values were extracted, we use sea surface values to model habitat suitability for planktivores, which are largely pelagic species, and seabed values for the benthivores and piscivores which are largely demersal species. Pairwise Pearson correlation coefficients were computed for the set of environmental variables used to model the habitat of planktivores and of non-planktivores separately, to assess multicollinearity. Variables were removed if they correlated with another &gt;0.7.&#13;
Additionally we provide data on model performance and presence/absence thresholds, in the file "Model performance and threshold.xlsx". Model performance is measured via the area under the curve (AUC) of the Receiver Operating Characteristic (ROC) plot, the AUC of the Precision-Recall (PR) plot (He &amp; Garcia, 2009), and the Miller slope (Miller et al., 1991). We assess model performance in both space and time. Spatial performance was analysed via 8-fold block cross validation using the R library “blockCV” (Valavi et al., 2019). Temporal performance of the models was assessed by training a new model that excluded the last 5 years of survey data (2015 to 2019), using the resulting model to predict to those 5 years, a “novel time period” for that model. &#13;
These data were produced for the study in the article "Climate change affects the distribution of diversity across marine food webs", by Murray S. A. Thompson, Elena Couce, Michaela Schratzberger and Christopher P. Lynam, in Global Change Biology (in press).</Abstract>
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&#13;
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        <Abstract>The dataset includes 6  layers showing modelled habitat suitability for harbour porpoises (Phocoena phocoena) across the North Sea–Baltic Sea transition zone over a 25-year period (1997–2004; 2005-2012; 2013-2022) during winter (Nov–Apr) and summer (May–Oct). Habitat suitability was estimated using the maximum entropy (MaxEnt) algorithm, based on satellite tag data of 111 individuals and a range of dynamic and static environmental conditions as predictor variables. Habitat suitability models and maps were developed by researchers of the Department of Ecoscience, Aarhus University to quantify potential spatiotemporal changes in the overlap between suitable habitats and an existing network of 62 MPAs dedicated to harbour porpoise protection. The habitat suitability rasters can inform effective design of forthcoming MPAs under the EU Biodiversity Strategy for 2030.</Abstract>
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        <Abstract>The large databases of EMODNET Biology only store confirmed presences of species. However, when mapping species distribution, it is also important where the species did not occur: there is at least as much information in absences as in presences. Inferring absences from presence-only databases is difficult and always involves some guesswork. In this product we have used as much meta-information as possible to guide us in inferring absences.</Abstract>
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&#13;
As many biodiversity estimates are biased by sampling effort, caution must be taken when interpreting alpha diversity from microscopy counts. By rarefaction and evenness estimation, these biases can be reduced, but not ignored.&#13;
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This data product provides a spatial-temporal interpolation of zooplankton alpha diversity in the greater Baltic Sea area (including Skagerrak and Kattegat). Using observed Shannon diversity indices from environmental monitoring data, the product employs the DIVAnd (Data-Interpolating Variational Analysis in n-Dimensions) to create a three-dimensional interpolation across longitude, latitude, and seasonal time dimensions.&#13;
&#13;
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